The genus
Aspects of
Specimens were examined with dissecting microscopes and are deposited at the following institutional collections: California Academy of Sciences, San Francisco California, United States of America (CAS); Canadian National Collection of Insects, Ottawa, Ontario, Canada (CNC); Cornell University Insect Collection, Ithaca, New York, United States of America (CUIC); Purdue University Entomological Research Collection, West Lafayette, Indiana, United States of America (PERC).
HOLOTYPE: United States of America, California, Madera County, Sierra National Forest, Chilacoot Creek, 1 mile above Bass Lake, 19-VI-1959, R. K. Allen, larva (PERC).
HOLOTYPE: United States of America, Oregon, Clackamas County, Branch of Still Creek on road to Timberline Lodge, Mt. Hood, 30-VIII-1958, G. F. Edmunds, Jr., & R. K. Allen, larva (PERC).
HOLOTYPE: United States of America, Oregon, Douglas County, Umpqua River, Sawyer Rapids, 26.5 mi E of Reedsport, 17-VI-1958, Mike Johnson, larva (PERC).
HOLOTYPE: Canada, British Columbia, Peachland, Trepanier Creek, 4-VII-1934, A. N. Gartrell, male adult (reared from larva) (CNC).
HOLOTYPE: United States of America, Wyoming, Yellowstone National Park, Upper Geyser Basin, 29-VII-1928, J. McDunnough, male adult (CNC).
NEOTYPE (new designation), United States of America, Montana, Sweet Grass County, small tributary of Big Timber Creek, at canyon road crossing, 1 mile south of Big Moon Campground, 46°1′30′′N, 110°8′48′′W, 10-VI-2000, W. P. McCafferty et al., larva (CUIC).
HOLOTYPE, United States of America, California, Inyo County, 4-VII-1938, larva (specimen number 8645) (CAS).
United States of America, California, Los Angeles County, San Gabriel River at Camp Bonita (62°), 16-VI-1959, R. K. Allen, one larva, paratype (PERC).
United States of America, Oregon, Clackamas County, Branch of Still Creek on road to Timberline Lodge, Mt. Hood, 30-VIII-1958, G. F. Edmunds, Jr., & R. K. Allen, two larvae (same collection data as holotype) (CNC); United States of America, Washington, Pierce County, Mt. Rainier National Park, spring-fed stream, Westside Road, 1.2 miles north of Highway 706, 16-VI-2004, emerged 21-VI, Kondratieff, Schmidt, three male adults, one female adult, associated larval exuviae (PERC).
Canada, British Columbia, Peachland, Trepanier Creek (same collection locale as holotype), A. N. Gartrell, sub 15-VII-1934, adult 17-VII-1934, one set larval exuviae (CNC); same locale and collector, sub 6-VII-1934, adult 10-VII-1934, one set larval exuviae (CNC); same locale and collector, sub 5-VII-1934, adult 6-VII-1934, one set larval exuviae (CNC).
United States of America, Wyoming, Yellowstone National Park, Firehole River, above Chimney Bridge, 22-VII-1928, J. McDunnough, nine larvae (CNC).
Canada, British Columbia, Keremos, Shingle Creek Road, sub 26-VII-1935, adult 28-VII-1935, A. N. Gartrell (699-1192) (CNC); United States of America, Montana, Sweet Grass County, small tributary of Big Timber Creek, at canyon road crossing, 1 mile south of Big Moon Campground, 46°1′30′′N, 110°8′48′′W, 10-VI-2000, W. P. McCafferty, et al., (same collection data as neotype), five larvae (PERC).
This species complex is defined herein to contain two nominal species. Adults and larvae of these two related species are distinguished from congeners by abdominal sterna that have three longitudinal stripes and by cerci that are much shorter than the median filament (usually less than half as long).
In contrast to
Male adults of
In contrast to
Male adults of
Allen and Edmunds’ [
This species complex consists of one polytypic species. Larvae are distinguished from congeners by having long, almost hook-like, paired medial spines on the middle abdominal terga. Adults are distinguished from congeners by having abdominal sterna with broad medial maculation that occupies most of the segment, usually taking the form of a chevron shape [
The name
In order to emphasize the polytypic nature of
The
Unfortunately, the type specimen (no. 1287.1) of
In order to maintain the identity of
The neotype larva of
The somewhat discolored
Previous study indicated that the maculation of larval sterna can be variable [
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Maxillary palp vestigial; tarsal claw with two prominent rows of denticles—(
Maxillary palp with three distinct segments; tarsal claw with only one distinct row of denticles—(2).
Paired medial spines on abdominal terga long and curved (at least on middle segments), some almost hook like—(
Paired medial spines on abdominal terga relatively straight, none longer than respective segment—(3).
Cerci approximately two thirds length of median filament; abdominal sterna with solid color, never with longitudinal stripes or other such markings—(
Cerci less than one-half length of median filament; abdominal sterna almost always with three dark, longitudinal markings—(4).
Cerci approximately one-third length of median filament (about the length of the abdomen); distinct pair of medial spines present only on abdominal terga 2–9, with the spine tips blunt—(
Cerci approximately one-sixth length of median filament (about half the length of the abdomen); distinct pair of medial spines present on abdominal terga 1–9, with the spine tips sharp—(
Richard Hoebeke (Cornell University, Ithaca, New York, USA), Vincent Lee (California Academy of Science, San Francisco, California, USA), Patrick McCafferty (Purdue University, West Lafayette, Indiana, USA), and Janice Peters and Bart Richard (Florida A&M University, Tallahassee, Florida, USA) assisted in attempting to locate the